Ecological interactions at the intersection of spatial ecology, global change, and behavioral ecology.


Channel Islands sampling (Credit: Raina Eddy)
Quantifying granivory (Credit: Jen Chandler)
Rodent foraging (Credit: John Orrock)


Connolly, B. M., C. Zirbel, and J. L. Orrock. In press. Rodent-mediated seed limitation affects woody seedling establishment more than invasive shrubs and downed woody debris.
Canadian Journal of Forest Research.

Lanni, F., Connolly, B. M., J. L. Orrock, and P. W. Guiden. 2024. Seed coatings containing capsaicin reduce seed removal of temperate woody species.
Canadian Journal of Forest Research 54:238-244. [Reprint]


Brehm, A. M. and J. L. Orrock. 2023. Extensive behavioral data contained within existing ecological datasets.
Trends in Ecology and Evolution 38:1129-1133. [Reprint]

Bartel, S. L., J. C. Kilgo, and J. L. Orrock. 2023. White-tailed deer responses to acoustic predator cues are contingent upon past land use and contemporary fire regime.
Behavioral Ecology 34:1013-1022. [Reprint]

Keller, C. B. and J. L. Orrock. 2023. Removal of invasive shrubs reduces rodent consumption of non-native invasive animals and native understory seeds.
Biological Invasions 25:3207-3221. [Reprint]

Orrock, J. L., L. A. Brudvig, E. I. Damschen, W. B. Mattingly, J. Cruz, J. W. Veldman, P. G. Hahn, and A. L. Larsen-Gray. 2023. Long-term, large-scale experiment reveals the profound effect of seed limitation, climate, and anthropogenic effects on the restoration of plant communities in a biodiversity hotspot.
Proceedings of the National Academy of Sciences USA 120: e2201943119. [Reprint]

Bartel, S. L. and J. L. Orrock. 2023. Land-use history, fire regime, and large-mammal herbivory affect deer-preferred plant diversity in longleaf pine woodlands.
Forest Ecology and Management 541:121023. [Reprint]

LoPresti, E., V. S. Pan, C. Girvin, G. Barber, S. Jaeger, and J. L. Orrock. 2023. Sandy seeds: armor or invisibility cloak? Mucilage-bound sand physically protects seeds from rodents and invertebrates.
Journal of Ecology 111:540-551. [Reprint]

Gilbert, N., K. McGinn, L. Nunes, A. Shipley, J. Bernath-Plaisted, J. Clare, P. Murphy, S. Keyser, K. Thompson, S. Maresh Nelson, J. Cohen, I. Widick, S. Bartel, J. L. Orrock, and B. Zuckerberg. 2023. Daily activity timing in the Anthropocene.
Trends in Ecology and Evolution 38:324-336. [Reprint]

Bartel, S. L., J. T. Schamel, B. A. Parrino, M. K. Marshall and J. L. Orrock. 2023. Channel Island foxes are viable seed dispersal agents of toyon.
Western North American Naturalist 83:23-32. [Reprint]

Resasco, J., M. Burt, J. L. Orrock, N. M. Haddad, D. Shoemaker, and D. J. Levey. 2023. Transient effects of corridors on polygyne fire ants over a decade.
Ecological Entomology 48:263-268. [Reprint]


Bartel, S. L. and J. L. Orrock. 2022. The important role of animal social status in vertebrate seed dispersal.
Ecology Letters 25:1094-1109. [Reprint]

Orrock, J. L., P. W. Guiden, V. S. Pan, and R. Karban. 2022. Plant induced defenses that promote cannibalism reduce herbivory as effectively as highly pathogenic herbivore pathogens.
Oecologia 199:397-405. [Reprint]

Anderegg, G. C, J. J. Henn, J. L. Orrock, E. I. Damschen. 2022. Litter removal reduces seed predation in restored prairies during times when seed predation would otherwise be high.
Restoration Ecology 30:e13550 [Reprint]


Bartel, S. L. and J. L. Orrock. 2021. Past agricultural land use affects multiple facets of ungulate antipredator behavior.
Behavioral Ecology 32:961-969. [Reprint]

Orrock, J. L., L. Abueg, S. Gammie, and J. Munshi-South. 2021. Exome sequencing of deer mice on two California Channel Islands identifies potential adaptation to strongly contrasting ecological conditions.
Ecology and Evolution 11:17191-17201. [Reprint]

Orrock, J. L., B. M. Connolly, P. W. Guiden, J. L. Chandler, G. M. Bron, C. A. Drost, and D. K. Garcelon. 2021. Sin Nombre virus prevalence in wild deer mice, Peromyscus maniculatus, on five of the California Channel Islands 2014-2017.
Zoonoses and Public Health 68:849-853. [Reprint]

Brudvig, L. A., N. E. Turley, S. L. Bartel, L. Bell-Dereske, S. Breland, E. I. Damschen, S. E. Evans, J. Gibbs, P. G. Hahn, R. Isaacs, J. A. Ledvina, J. L. Orrock, Q. Sorenson, J. D. Stuhler. 2021. Large ecosystem-scale effects of restoration fail to mitigate impacts of land-use legacies in longleaf pine savannas.
Proceedings of the National Academy of Sciences USA 118:e2020935118. [Reprint]

Bartel, S. and J. L. Orrock. 2021. An omnivorous mesopredator modifies predation of omnivore-dispersed seeds.
Ecosphere 12:e03369. 10.1002/ecs2.3369. [Reprint]

Guiden, P. W. and J. L. Orrock. 2021. Snow depth and woody debris drive variation in small mammal winter seed removal.
Journal of Vegetation Science 32:e13007. [Reprint]

Orrock, J. L. 2021. A simple method for restraint of small mammals for sampling blood or tissue in the field.
Western North American Naturalist 81:264-266. [Reprint]


Sheriff, M., J. L. Orrock, M. Ferrari, R. Karban, E. Preisser, A. Sih, and J. Thaler. 2020. Proportional fitness loss and the timing of defensive investment: a cohesive framework across animals and plants.
Oecologia 193:273-283. [Reprint]

Chandler, J. L., T. Van Deelen, N.. Nibbelink, and J. L. Orrock. 2020. Large-scale patterns of seed removal by small mammals differ between areas of low- vs. high wolf occupancy.
Ecology and Evolution 10:7145-7156. [Reprint]

Bartel, S.L. and J. L. Orrock. 2020. Past and present disturbances generate spatial variation in seed predation.
Ecosphere 11:e03116. [Reprint]

Orrock, J. L. 2020. Deterministic insights from stochastic interactions.
Proceedings of the National Academy of Sciences USA 117:6965-6957. [Reprint]

Guiden, P. W. and J. L. Orrock. 2020. Seasonal shifts in activity timing reduce small-mammal heat loss during winter.
Animal Behaviour 164: 181-192. [Reprint]

Raffa, K., P. Bonello, and J. L. Orrock. 2020. Why do entomologists and plant pathologists approach trophic relationships so differently? Identifying biological distinctions to foster synthesis.
New Phytologist 225:609-620. [Reprint]

Donelan, S. C., J. K. Hellmann, A. M. Bell, B. Luttbeg, J. L. Orrock, M. J. Sheriff, and A. Sih. 2020. Transgenerational plasticity in human-altered environments.
Trends in Ecology and Evolution 35:115-124. [Reprint]

Connolly, B. M., P. W. Guiden, and J. L. Orrock. 2020. Mycorrhizal inoculation mitigates damage from an intermediate, but not severe, frost event for a cool-season perennial bunchgrass.
Botany 98:127-135.  [Reprint]


Damschen, E. I., L. A Brudvig, M. A. Burt, R. J. Fletcher, Jr., N. M. Haddad, D. J. Levey, J. L. Orrock, J. Resasco, J. J. Tewksbury. 2019. Landscape connectivity causes diversity to accumulate over decades through colonization credits and reduced extinctions.
Science 365:1478-1480. [Reprint]

Guiden, P. W., S. L. Bartel, N. W. Byer, A. A. Shipley, and J. L. Orrock. 2019. Predator-prey interactions in the Anthropocene: reconciling multiple aspects of novelty.
Trends in Ecology and Evolution 34:616-627. [Reprint]

Kelt, D. A., E. J. Heske, X. Lambin, M. K. Oli, J. L. Orrock, A. Ozgul, J. N. Pauli, L. Prugh, R. Sollmann, and S. Sommer. 2019. Advances in population ecology and species interactions in mammals.
Journal of Mammalogy 100: 965-1007. [Reprint]

Guiden, P. W. and J. L. Orrock. 2019. Invasive shrubs modify rodent activity timing, revealing a consistent behavioral rule governing diel activity.
Behavioral Ecology: 30:1069-1075. [Reprint]

Barker, C., Turley, N., J. L. Orrock, J. Ledvina, and L A. Brudvig. 2019. Agricultural land-use history does not reduce woodland understory herb establishment.
Oecologia 189: 1049-1060. [Reprint]

Guiden, P. W., B. M. Connolly, and J. L. Orrock. 2019. Seedling responses to decreased snow depend on canopy composition and small-mammal herbivore presence.
Ecography 42:780-790. [Reprint]

Pearson, D. E., M. Valliant, C. Carlson, G. C. Thelen, Y. K. Ortega, J. L. Orrock, and M. D. Madsen. 2019. Spicing up restoration: can chili peppers improve reseeding success by reducing seed predation?
Restoration Ecology 27:254-260. [Reprint]


Karban, R. and J. L. Orrock. 2018. A judgement and decision-making model for plant behavior.
Ecology 99:1909-1919. [Reprint]

Orrock, J. L., B. M. Connolly, W.-G. Choi, P. W. Guiden, S. W. Swanson, and S. Gilroy. 2018. Plants eavesdrop on cues produced by snails and induce costly defenses that affect insect herbivores.
Oecologia 186:703-710. [Reprint]

Sheriff, M. J., B. Dantzer, O. P. Love, and J. L. Orrock. 2018. Error management theory and the adaptive significance of transgenerational maternal-stress effects on offspring phenotype.
Ecology and Evolution 8:6473-6482. [Reprint]

Connolly, B. M. and J. L. Orrock. 2018. Exogenous application of methyl jasmonate alters Pinus resinosa seedling response to simulated frost.
Botany 96:705-710. [Reprint]

Connolly, B. M., J. L. Orrock. 2018. Habitat-specific capture timing of deer mice (Peromyscus maniculatus) suggests that predators structure temporal activity of prey.
Ethology 124:105-112. [Reprint]

Guiden, P. W., B. M. Connolly, and J. L. Orrock. 2018. Extreme cold consistently reduces seedling growth but has species-specific effects on browse tolerance in summer.
American Journal of Botany 105:2075-2080. [Reprint]

Schwemm, C. A., C. A. Drost, T. J. Coonan, J. L. Orrock, and T. R. Stanley. 2018. Comparison of density estimators for monitoring long-term population trends in deer mice, Peromyscus maniculatus, on the California Channel Islands.
Western North American Naturalist 78:496-509.


Orrock, J. L., B. M. Connolly, and A. J. Kitchen. 2017. Plant induced defences reduce herbivory by increasing cannibalism.
Nature Ecology and Evolution 1:1205-1207. [Reprint]

Connolly, B. M., P. W. Guiden, and J. L. Orrock. 2017. Past freeze-thaw events on Pinus seeds increase seedling herbivory.
Ecosphere 8(3):e01748. [Reprint]

Turley, N., J. L. Orrock, J. Ledvina, and L. Brudvig. 2017. Dispersal and establishment limitation slows plant community recovery on post-agricultural longleaf pine savannas.
Journal of Applied Ecology 54:1100-1109. [Reprint]

Connolly, B. M., L. Agnew, and J. L. Orrock. 2017. Interactive effects of fungicide and cold stratification on the germination rate for five dominant temperate tree species.
Forest Science 63:303-309. [Reprint]

Guiden, P. W. and J. L. Orrock. 2017. Invasive shrub modifies a classic animal-habitat relationship and alters patterns of vertebrate seed predation.
Ecology 98:321-327. [Reprint]

Collins, C., C. Banks-Leite, L. Brudvig, B. Foster, W. Cook, E. Damschen, A. Andrade, M. Austin, J. Camargo, D. Driscoll, R. Holt, W. Laurance, N. Nicholls, and J. L. Orrock. 2017. Fragmentation affects plant community composition over time.
Ecography 40:119-130. [Reprint]


Karban, R., J. L. Orrock, E. L. Preisser, and A. Sih. 2016. A comparison of plants and animals in their responses of risk of consumption.
Current Opinion in Plant Biology 32:1-8. [Reprint]

Hahn, P. G. and J. L. Orrock. 2016. Ontogenetic responses of four plant species to interactive effects of land-use history, canopy structure, and herbivory.
Journal of Ecology 104: 1518-1526. [Reprint]

Orrock, J. L. and Connolly, B. M. 2016. Changes in trap temperature as a method to determine timing of activity of small mammals.
PLOS ONE 11(10):e0165710. [Reprint]

Connolly, B. M., J. L. Orrock, and M. S. Witter. 2016. Soil conditions mediate the effect of herbivores, but not mycorrhizae, on a native bunchgrass.
Acta Oecologica 77: 100-108. [Reprint]

Hahn, P. G. and J. L. Orrock. 2016. Neighbor palatability generates associational effects by altering herbivore foraging behavior.
Ecology 104:1518-1526. [Reprint]

Grace, J. B., T. M. Anderson, E. W. Seabloom, E. T. Borer, P. B. Adler, W. S. Harpole, Y. Hautier, H. Hillebrand, E. M. Lind, M. Partel, J. D. Bakker …, J. L. Orrock, S. M. Prober, and M. D. Smith [26 authors total]. 2016. Integrative modeling reveals mechanisms linking productivity and plant species richness.
Nature 529:390-393. [Reprint]

Chandler, J. L., J. L. Orrock, and J. Resasco. 2016. Invasive ants generate heterogeneity in patterns of seed survival.
American Midland Naturalist 176: 289-297. [Reprint]

Stuhler, J. D. and J. L. Orrock. 2016. Historical agriculture and present-day canopy thinning differentially affect the distribution and abundance of invasive and native ant species.
Biological Invasions 18:1813-1825. [Reprint]

Stuhler, J. D. and J. L. Orrock. 2016. Past agricultural land use and present-day fire regimes can interact to determine the nature of seed predation.
Oecologia 181:463-473. [Reprint]

Herrmann, J. D., T. A. Carlo, L. A. Brudvig, E. I. Damschen, N. M. Haddad, D. J. Levey, J. L. Orrock, and J. J. Tewksbury. 2016. Connectivity from a different perspective: comparing seed dispersal kernels in connected vs. unfragmented landscapes.
Ecology 97:1274-1282. [Reprint]

Bartowitz, K. J. and J. L. Orrock. 2016. Invasive exotic shrub (Rhamnus cathartica) alters the timing and magnitude of post-dispersal seed predation of native and exotic species.
Journal of Vegetation Science 27:789-799. [Reprint]


Orrock, J. L., A. Sih, M. C. O. Ferrari, R. Karban, E. L. Preisser, M. J. Sheriff, and J. Thaler. 2015. Error management in plant allocation to herbivore defense.
Trends in Ecology and Evolution 30:441-445. [Reprint]

Orrock, J. L., H. P. Dutra, R. J. Marquis, and N. Barber. 2015. Apparent competition and native consumers exacerbate the strong competitive effect of an exotic plant.
Ecology 96:1052-1061. [Reprint]

Connolly, B. M. and J. L. Orrock. 2015. Climatic variation virtually eliminates seed persistence: freeze-thaw cycles reduce seed survival via the joint action of abiotic stress and fungal pathogens.
Oecologia 179:609-616. [Reprint]

Hahn, P. G. and J. L. Orrock. 2015. Land-use history alters contemporary insect herbivore community composition and decouples plant-herbivore relationships.
Journal of Animal Ecology 84:745-754. [Reprint]

Hahn, P. G. and J. L. Orrock. 2015. Spatial arrangement of canopy structure and land-use history alter herbivory in a landscape experiment.
Ecosphere 6:193. [Reprint]

Grman, E., J. L. Orrock, C. Habeck, J. Ledvina, L. Brudvig. 2015. Altered beta diversity in post-agricultural woodlands: two hypotheses and the role of scale.
Ecography 38:614-621. [Reprint]

Orrock, J. L., E. T. Borer, L. A. Brudvig, J. Firn, A. S. MacDougall, B. A. Melbourne, L. H. Yang, D. V. Baker P, A. Bar-Massada, M. J. Crawley, E. I. Damschen, K. F. Davies, D. S. Gruner, A.D. Kay, E. Lind, R. L. McCulley, E. W. Seabloom.  2015. A continent-wide study reveals clear relationships between regional abiotic conditions and post-dispersal seed predation.
Journal of Biogeography 42:662-670. [Reprint]

Hahn, P. G. and J. L. Orrock. 2015. Land-use legacies and present fire regimes interact to mediate herbivory by altering the neighboring plant community.
Oikos 124:497-506. [Reprint]

Haddad, N. M., L. A. Brudvig, J. Clobert, K. F. Davies, A. Gonzalez, R. D. Holt, T. E. Lovejoy, J. O. Sexton, M. P. Austin…, J. L. Orrock, D-X Song, and J. R. Townshend [24 authors total]. 2015. Habitat fragmentation and its lasting impact on Earth’s ecosystems.
Science Advances 2015: e1500052. [Reprint]

Seabloom, E. S., E. T. Borer, Y. Buckley, E. Cleland, K. Davies, J. Firn, W. S. Harpole Y. Hautier, E. Lind, A. MacDougall,J. L. Orrock, S. M. Prober, P. Adler, T. M. Anderson…, J.Wright, L. Yang [60 authors total]. 2015. Plant species’ origin determines dominance and response to nutrient enrichment and herbivores.
Nature Communications 6: 7710. [Reprint]

Tredennick, A. T., P. B. Adler, J. B. Grace, W. S. Harpole, E. T. Borer, E. W. Seabloom, T. M. Anderson, J. D. Bakker, … J. L. Orrock, …, R. J. Williams, L. Yang [41 authors total]. 2015. Comment on “Worldwide evidence of a unimodal relationship between productivity and plant species richness.
Science 335: 1441. [Reprint]

Stevens, C. J., E. M. Lind, Y. Hautier, W. S. Harpole, E. T. Borer, S. Hobbie, E. W. Seabloom …, J. L. Orrock, S. M. Prober, A. Risch, M. Schultz, and P. D. Wragg [24 authors total]. 2015. Anthropogenic nitrogen deposition predicts local grassland primary production worldwide.
Ecology 96:1459-1465. [Reprint]

Mattingly, W. B., J. L. Orrock, C. D. Collins, L. A. Brudvig, E. I. Damschen, J. W. Veldman, and J. L. Walker. 2015. Historical agriculture alters the effects of fire on understory plant beta diversity.
Oecologia 177:507-518. [Reprint]


Veldman, J. W., L. A. Brudvig, E. I. Damschen, J. L. Orrock, W. B. Mattingly, and J. L. Walker.  2014. Fire frequency, agricultural history, and the multivariate control of pine savanna understory plant diversity.
Journal of Vegetation Science 25:1438-1449. [Reprint]

Orrock, J. L., and R. J. Fletcher, Jr. 2014. An island-wide predator manipulation reveals immediate and long-lasting matching of risk by prey.
Proceedings of the Royal Society B: 281:20140391. [Reprint]

Haddad, N. M., L. A. Brudvig, E. I. Damschen, D. M. Evans, B. L. Johnson, D. J. Levey, J. L. Orrock, J. Resasco, L. L. Sullivan, J. J. Tewksbury, S. A. Wagner, and A. J. Weldon. 2014. A review of the potential negative ecological effects of landscape corridors.
Conservation Biology 28:1178-1187. [Reprint]

MacDougall, A. S., J. R. Bennett, J. L. Firn, Seabloom, E. S., E. T. Borer, E. M. Lind, J. L. Orrock, ….J. W. Morgan, and C. S. Stevens [22 authors total]. 2014. Regional-scale factors most consistently explain plot-level diversity in invaded areas.
Global Ecology and Biogeography 23:802-810. [Reprint]

Resasco, J., N. M. Haddad, J. L. Orrock, D. Shoemaker, L. A. Brudvig, E. I. Damschen, J. J. Tewksbury, and D. L. Levey. 2014. Landscape corridors can increase invasion by an exotic species and reduce diversity of native species.
Ecology 95:2033-2039. [Reprint]

Borer, E., T., E. S. Seabloom, D. Gruner, W. S. Harpole, H. Hillebrand, E. Lind, P. Adler, … J. L. Orrock, …, J.Wright, L. Yang [55 authors total].  2014. Herbivores and nutrients control grassland plant diversity via light limitation.
Nature 508:517-520. [Reprint]

Damschen, E. I., D. V. Baker P, G. Bohrer, R. Nathan, J. L. Orrock, J. R. Turner, L. A. Brudvig, N. M. Haddad, D. J. Levey, and J. J. Tewksbury.  2014. Models and experiments reveal wind-driven seed dispersal is affected by fragmentation and corridors.
Proceedings of the National Academy of Sciences USA 111:3484-3489. [Reprint]

Brudvig, L. A., J. L. Orrock, E. I. Damschen, C. Collins, P. G. Hahn, W. B. Mattingly, J. W. Veldman, and J. L. Walker. 2014. Land-use history and contemporary management inform an ecological reference model for longleaf pine woodland understory plant communities.
PLOS ONE 9:e86604. [Reprint]

Borer, E. T., W. S. Harpole, P. B. Adler, E. M. Lind, J. L. Orrock, E. W. Seabloom, and M. D. Smith.  2014. Finding generality in ecology: a model for globally-distributed experiments.
Methods in Ecology and Evolution 5:65-73. [Reprint]

Hahn, P. G. and J. L. Orrock. 2014. Effects of temperature on seed viability of six Ozark glade herb species and eastern red cedar (Juniperus virginiana).
American Midland Naturalist 171:147-152. [Reprint]


Orrock, J. L. 2013. Exposure of unwounded plants to chemical cues associated with herbivores leads to exposure-dependent changes in subsequent herbivore attack.
PLOS ONE 8:e79900. [Reprint]

Seabloom, E. S., E. T. Borer, Y. Buckley, E. Cleland, K. Davies, J. Firn, W. S. Harpole Y. Hautier, E. Lind, A. MacDougall, J. L. Orrock, S. M. Prober, P. Adler, T. M. Anderson…, J.Wright, and L. Yang [60 authors total]. 2013. Dominance by invasive species is the real embarrassment of riches: invasion in grassland ecosystems.
Global Change Biology 19:3677-3687. [Reprint]

Brudvig, L. A., E. Grman, C. Habeck, J. L. Orrock, and J. Ledvina.  2013. Strong legacy of agricultural land use on soils and understory plant communities in longleaf pine woodlands.
Forest Ecology and Management 310:944-955. [Reprint]

Mattingly, W. B. and J. L. Orrock.  2013. Historic land use influences contemporary establishment of invasive plant species.
Oecologia 172:1147-1157. [Reprint]

Orrock, J. L., E. L. Preisser, J. H. Grabowski, and G. C. Trussell. 2013. The cost of safety for prey: refuges increase the impact of predation risk in aquatic systems.
Ecology 94:573-579. [Reprint]

Coyle, D. R., M. M. Murphy, S. M. Paskewitz, J. L. Orrock, X. Lee, R. J. Murphy, M. A. McGeehin, and K. F. Raffa.  2013.  Belowground herbivory in red pine stands initiates a cascade that increases abundance of Lyme disease vectors.
Forest Ecology and Management 302:354-362. [Reprint]

Mattos, K. J., J. L. Orrock, and J. I. Watling. 2013. Rodent granivores generate context-specific seed removal in invaded and uninvaded habitats.
American Midland Naturalist 169:168-178. [Reprint]

O’Halloran, L. R., E. T. Borer, E. W. Seabloom, A. S. MacDougall, E. E. Cleland, R. L. McCulley, S. Hobbie, W. S. Harpole, N. M. DeCrappeo, C. Chu, J. D. Bakker, K. F. Davies, G. Du, J. Firn, N. Hagenah, K. S. Hofmockel, J. M. H. Knops, W. Li, B. A. Melbourne, J. W. Morgan, J. L. Orrock, S. M. Prober, and C. J. Stevens. 2013. Regional contingencies in the relationship between aboveground biomass and litter in the world’s grasslands.
PLoS ONE 8:e54988. [Reprint]


Mattingly, W.B., J. L. Orrock, and N. T. Reif. 2012. Dendroecological analysis reveals long-term, positive effects of an introduced understory plant on canopy tree growth.
Biological Invasions 14:2639-2646. [Reprint]

Fletcher, R. J. Jr., J. L. Orrock, and B. A. Robertson. 2012.  How the type of anthropogenic change alters the consequences of ecological traps.
Proceedings of the Royal Society B 279:2456-2552. [Reprint]

Preisser, E. L. and J. L. Orrock. 2012. The allometry of fear: Interspecific relationships between body size and response to predation risk.
Ecosphere 3:77. [Reprint]

Schuler, M. S. and J. L. Orrock. 2012. The maladaptive significance of maternal effects for plants in anthropogenically modified environments.
Evolutionary Ecology 26:475-481. [Reprint]

Orrock, J. L., C. C. Christopher, and H. P. Dutra.  2012. Seed bank survival of an invasive species, but not two native species, declines with invasion.
Oecologia 168:1103-1110. [Reprint]

Grace, J. B., P. B. Adler, E. W. Seabloom, E. T. Borer, H. H. Hillebrand, Y. Hautier, A. Hector, W. S. Harpole, L. R. O’Halloran, …J. L. Orrock… ,L. H. Yang [37 authors total].  2012. Response to Comments on ” Productivity is a poor predictor of plant species richness”: towards a multivariate representation of the multiple mechanisms controlling productivity and diversity.
Science 335: 1441.


Adler, P. B., E. W. Seabloom, E. T. Borer, H. H. Hillebrand, Y. Hautier, A. Hector, W. S. Harpole, L. R. O’Halloran, J. B. Grace, …J. L. Orrock… ,L. H. Yang [58 authors total].  2011. Productivity is a poor predictor of plant species richness.
Science 333:1750-1753. [Reprint]

Orrock, J. L., G. R. Curler, B. J. Danielson, and D. R. Coyle.  2011. Large-scale experimental landscapes reveal distinctive effects of patch shape and connectivity on arthropod communities.
Landscape Ecology 26:1361-1372. [Reprint]

Craig, M. T., J. L. Orrock, and L. A. Brudvig. 2011. Edge-mediated patterns of seed removal in experimentally connected and fragmented landscapes.
Landscape Ecology 26:1373-1381.[Reprint]

Hinkelman, T. M., J. L. Orrock, and S. C. Loeb. 2011. Effect of downed woody debris on small mammal anti-predator behavior.
Ethology 118:17-23. [e-mail me for a reprint]

Watling, J. I., C. R. Hickman, and J. L. Orrock. 2011. Invasive shrub alters native forest amphibian communities.
Biological Conservation 144: 2597-2601. [Reprint]

Mulder, C. P. H., H. Jones, K. Kameda, C. Palmborg, S. Schmidt, J. Ellis, J. L. Orrock, A. Wait, D. A. Wardle, L. Yang, H. Young G, D. Croll, and E. Vidal. 2011. Impacts of seabirds on plant and soil properties. In: Seabird islands: Ecology, Invasions, and Restoration. C. P. H. Mulder, W. B. Anderson, D. R. Towns, and P. J. Bellingham, editors.  Oxford University Press.

Haddad, N. M., B. Hudgens, E. I. Damschen, D. Levey, J. L. Orrock, J. Tewksbury, and A. Weldon. 2011. Assessing both positive and negative effects of corridors. Pages 475-503 in: Sources, Sinks, and Sustainability. J. Liu, V. Hull, A. T. Morzillo, and J. A. Wiens, editors. Cambridge University Press.

Orrock, J. L., B. F. Allan, and C. A. Drost. 2011. Biogeographic and ecological regulation of disease: island area, precipitation, and predator richness are related to prevelance of Sin Nombre virus in island mice.
American Naturalist 177:691-697. [Reprint]

Dutra, H. P., K. Barnett, J. R. Reinhardt, R. J. Marquis, and J. L. Orrock. 2011. Invasive plant species alters consumer behavior by providing refuge from predation.
Oecologia 166:649-657. [Reprint]

Conley, A. K., Watling, J. I., and J. L. Orrock. 2011. Invasive plant alters ability to predict disease vector distribution.
Ecological Applications 21:329-334. [Reprint]

Watling, J. I., C. R. Hickman, and J. L. Orrock. 2011. Predators and invasive plants affect performance of amphibian larvae.
Oikos 120: 735-739. [Reprint]

Watling, J. I., A. J. Nowakowski, M. A. Donnelly, and J. L. Orrock. 2011. Meta-analysis reveals the importance of matrix composition for animals in fragmented habitat.
Global Ecology and Biogeography 20:209-217. [Reprint]

Watling, J. I., C. R. Hickman, E. Lee, K. Wang, and J. L. Orrock. 2011. Extracts of the invasive shrub Lonicera maackiiincrease mortality and alter behavior of amphibian larvae.
Oecologia 165:153-159. [Reprint]


Allan, B. F., H. P. Dutra, L. S. Goessling, K. Barnett, J. M. Chase, R. J. Marquis, G. Pang, G. A. Storch, R. E. Thach, and J. L. Orrock. 2010. Invasive honeysuckle eradication reduces tick-borne disease risk by altering host dynamics.
Proceedings of the National Academy of Sciences USA 107:18523-18527. [Reprint]

Orrock, J. L., M. L. Baskett, and R. D. Holt. 2010. Spatial interplay of plant competition and consumer foraging mediate plant coexistence and drive the invasion ratchet.
Proceedings of the Royal Society B 277: 3307-3315. [Reprint]

Orrock, J. L. and J. I. Watling. 2010. Local community size mediates ecological drift and competition in metacommunities.
Proceedings of the Royal Society B 277: 2185-2191. [Reprint]

Mattos, K. J., and J. L. Orrock. 2010. Behavioral consequences of plant invasion: an invasive plant alters rodent anti-predator behavior.
Behavioral Ecology 21:556-561. [Reprint]

Orrock, J. L., L. Dill, A. Sih, J. Grabowski, S. Peacor, B. Peckarsky, E. Preisser, J. R. Vonesh and E. Werner. 2010. Predator effects in predator-free space: the remote effects of predators on prey.
The Open Ecology Journal 3:22-30 [Reprint]

Sih, A., D. I. Bolnick, B. Luttbeg, J. L. Orrock, S. D. Peacor, L. M. Pintor, E. Preisser, J. S. Rehage, and J. R. Vonesh. 2010. Predator-prey naivete, antipredator behavior, and the ecology of predator invasions.
Oikos 119:610-621. [Reprint]

Orrock, J. L. 2010. When the ghost of predation has passed: do rodents from islands with and without fox predators exhibit aversion to fox cues?
Ethology 116:338-345. [e-mail me for a reprint]

Orrock, J. L. and C. C. Christopher. 2010. Density of intraspecific competitors determines the occurrence and benefits of accelerated germination.
American Journal of Botany 97:694-699. [Reprint]

Orrock, J. L., R.D. Holt, and M. L. Baskett. 2010. Refuge-mediated apparent competition in plant-consumer interactions.
Ecology Letters 13:11-20. [Reprint]

Orrock, J. L. and M. S. Witter. 2010. Multiple drivers of apparent competition reduce re-establishment of a native plant in invaded habitats.
Oikos 119:101-108. [Reprint]

Watling, J. I. and J. L. Orrock. 2010. Measuring edge contrast using biotic criteria helps define edge effects on the density of an invasive plant.
Landscape Ecology 25:69-78. [Reprint]

Motheral, S. M., and J. L. Orrock.  2010. Gastropod herbivore preference for seedlings of two native and two exotic grass species.
American Midland Naturalist 163:106-114. [Reprint]

Heldermon, C. D., K. K. Ohlemiller, E. D. Herzog, C. Vogler, E. Y. Qin, D. F. Wozniak, Y. Tan, J. L. Orrock, and M. S. Sands. 2010. Therapeutic efficacy of bone marrow transplant, intracranial AAV-mediated gene therapy or both in the mouse model of MPS IIIB.
Molecular Therapy 18:873-880. [Reprint]


Orrock, J. L. and B. J. Danielson.  2009. Temperature and cloud cover, but not predator urine, affect winter foraging of mice.
Ethology 115:641-648. [e-mail me for a reprint]

Orrock, J. L. and J. L. Hoisington-López.  2009. Mortality of exotic and native seeds in invaded and uninvaded habitats.
Acta Oecologica 35: 758-262. [e-mail me for a reprint]

Seabloom, E. W., E. T. Borer, B. A. Martin, and J. L. Orrock.  2009. Effects of long-term consumer manipulations on invasion in oak savannah communities.
Ecology 90: 1356-1365. [Reprint]

Vellend, M. and J. L. Orrock.  2009. Ecological and genetic models of diversity: Lessons across disciplines. Pages 439-461 in: The Theory of Island Biogeography Revisited. J. Losos and R. E. Ricklefs, editors. Princeton University Press.

Orrock, J. L., M. S. Witter, and O. J. Reichman.  2009.  Native consumers and seed limitation constrain the restoration of a native perennial grass in exotic habitats.
Restoration Ecology 17: 148-157. [Reprint]


Damschen, E. I., L. A. Brudvig, N. M. Haddad, D. J. Levey, J. L. Orrock, and J. J. Tewksbury.  2008.  The movement ecology and dynamics of plant communities in fragmented landscapes.
Proceedings of the National Academy of Sciences USA 105: 19078-19083. [Reprint]

Orrock, J. L., J. H. Grabowski, S. D. Peacor, B. L. Peckarsky, E. L. Preisser, A. Sih, and E. E. Werner.  2008.  Consumptive and non-consumptive effects of predators on metacommunities of competing prey.
Ecology 89: 2426-2435. [Reprint]

Peckarsky, B. L., D. I. Bolnick, L. M. Dill, J. H. Grabowski, B. Luttbeg, J. L. Orrock, S. D. Peacor, E. L. Preisser, O. J. Schmitz, and G. C. Trussell.  2008. Revisiting the classics: Considering non-consumptive effects in textbook examples of predator-prey interactions.
Ecology 89: 2416-2425. [Reprint]

Orrock, J. L., M. S. Witter, and O. J. Reichman.  2008.  Apparent competition with an exotic plant reduces native plant establishment.
Ecology 89: 1168-1174. [Reprint]

Orrock, J. L. and B. A. Allan.  2008. Sin Nombre virus infection in deer mice, Channel Islands, California.
Emerging Infectious Diseases 14: 1965-1966. [Reprint]


Preisser, E. L., J. L. Orrock, and O. J. Schmitz.  2007.  Predator hunting mode and habitat domain affect the strength of non-consumptive effects in predator-prey interactions.
Ecology 88: 2744-2751. [Reprint]

Orrock, J. L. and E. I. Damschen.  2007.  The effect of burial depth on removal of seeds of Phytolacca americana.
Southeastern Naturalist 6: 151-158. [Reprint]


Orrock, J. L. 2006.  Useful distraction: Ritualized behavior as an opportunity for recalibration.
Behavioral and Brain Sciences 29: 625-626. [Reprint]

Orrock, J. L., D. J. Levey, B. J. Danielson, and E. I. Damschen.  2006.  Seed predation, not seed dispersal, explains the landscape-level abundance of an early-successional plant.
Journal of Ecology 94: 838-845. [Reprint]

Damschen, E. I., N. M. Haddad, J. L. Orrock, D. J. Levey, and J. J. Tewksbury.  2006.  Corridors increase plant species richness at large scales.
Science 313: 1284-1286. [Reprint]


Orrock, J. L. and R. J. Fletcher, Jr. 2005. Changes in community size affect the outcome of competition.
American Naturalist 166: 107-111. [Reprint]

Orrock, J. L.  2005. Conservation corridors affect the fixation of novel alleles.
Conservation Genetics 6: 623-630. [Reprint]

Orrock, J. L. and E. I. Damschen.  2005. Corridors cause differential seed predation.
Ecological Applications 15: 793-798. [Reprint]

Orrock, J. L. and B. J. Danielson. 2005. Patch shape, connectivity, and foraging by the oldfield mouse, Peromyscuspolionotus.
Journal of Mammalogy 86: 569-575. [Reprint]

Orrock, J. L.  2005. The effect of gut passage by two species of avian frugivores on seeds of Pokeweed, Phytolaccaamericana.
Canadian Journal of Botany 83: 427-431. [Reprint]

Orrock, J. L. and E. I. Damschen.  2005. Fungi-mediated mortality of seeds of two oldfield plant species.
Journal of the Torrey Botanical Society 132: 613-617. [Reprint]

Orrock, J. L. and B. J. Danielson.  2005. A note on the status of the endangered red-backed vole, Clethrionomys gapperi, in Iowa.
Journal of the Iowa Academy of Science 112: 24-25. [Reprint]

Brinkerhoff, R. J., N. M. Haddad, and J. L. Orrock.  2005. Corridors and olfactory predator cues affect small mammal behavior.
Journal of Mammalogy 86: 662-669. [Reprint]

2004 and earlier

Orrock, J. L. and B. J. Danielson.  2004. Rodents balancing a variety of risks: invasive fire ants and indirect and direct indicators of predation risk.
Oecologia 140: 662-667. [Reprint]

Orrock, J. L., B. J. Danielson, and J. Brinkerhoff.  2004. Rodent foraging is affected by indirect, but not by direct, cues of predation risk.
Behavioral Ecology 15: 433-437. [Reprint]

Orrock, J. L., B. J. Danielson, M. J. Burns, and D. J. Levey. 2003. Spatial ecology of predator-prey interactions:  corridors and patch shape influence seed predation.
Ecology 84: 2589-2599. [Reprint]

Orrock, J. L., D. Farley, and J. F. Pagels.  2003. Does fungus consumption by the woodland jumping mouse vary with habitat or the abundance of other small mammals?
Canadian Journal of Zoology 81: 753-756. [Reprint]

McShea, W. J., J. F. Pagels, J. L. Orrock, E. Harper, and K. Koy.  2003. Mesic deciduous forest as patches of small mammal richness within an Appalachian mountain forest.
Journal of Mammalogy 84: 627-643. [Reprint]

Orrock, J. L. and J. F. Pagels.  2003. Tree communities, microhabitat characteristics, and small mammals associated with the endangered rock vole, Microtus chrotorrhinus, in Virginia.
Southeastern Naturalist 2: 547-558. [Reprint]

Tewksbury, J. J., D. J. Levey, N. M. Haddad, S. Sargent, J. L. Orrock, A. Weldon, B. J. Danielson, J. Brinkerhoff, E. I. Damschen, and P. Townsend.  2002.  Corridors affect plants, animals, and their interactions in fragmented landscapes.
Proceedings of the National Academy of Sciences USA 99: 12923-12926. [Reprint]

Orrock, J. L. and J.F. Pagels.  2002. Fungus consumption by the southern red-backed vole, Clethrionomys gapperi, in the southern Appalachians.
American Midland Naturalist 147: 413-418. [Reprint]

Orrock, J. L., J.F. Pagels, W.J. McShea, and E.K. Harper.  2000. Predicting presence and abundance of a small mammal species: the effect of scale and resolution.
Ecological Applications 10: 1356-1366. [Reprint]

Orrock, J. L., E.K. Harper, J.F. Pagels, and W.J. McShea.  1999.  Additional records of the rock vole, Microtuschrotorrhinus (Mammalia: Muridae), in Virginia.
Banisteria: Journal of the Virginia Natural History Society 14: 36-38. [Reprint]